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"variegation" Definitions
  1. the act of variegating : the state of being variegated

83 Sentences With "variegation"

How to use variegation in a sentence? Find typical usage patterns (collocations)/phrases/context for "variegation" and check conjugation/comparative form for "variegation". Mastering all the usages of "variegation" from sentence examples published by news publications.

The pink splotches, called variegation, come from a genetic mutation.
Many cultivars of ivy can be found in and around gardens, some showing flashy variegation, others with finely dissected leaves.
The Rockies may offer wilder wildernesses, but you don't experience the pleasure of sharp cultural variegation as you move from place to place.
But these shifts feel deliberate, like the clicking of a slide projector rather than the smooth variegation of a kaleidoscope, and the landscape always clears atop that steady pulse.
Variegation in alt= Reflective variegation in Pilea cadierei. Some variegation is due to visual effects caused by reflection of light from the leaf surface. This can happen when an air layer is located just under the epidermis resulting in a white or silvery reflection. It is sometimes called blister variegation.
Some years there is more variegation, while others less. All-white branches are possible, but often die back. Otherwise, even with heavy variegation, the plant is very robust.
Because the variegation is due to the presence of two kinds of plant tissue, propagating the plant must be by a vegetative method of propagation that preserves both types of tissue in relation to each other. Typically, stem cuttings, bud and stem grafting, and other propagation methods that results in growth from leaf axil buds will preserve variegation. Cuttings with complete variegation may be difficult if not impossible to propagate. Root cuttings will not usually preserve variegation, since the new stem tissue is derived from a particular tissue type within the root.
Sometimes venal variegation occurs – the veins of the leaf are picked out in white or yellow. This is due to lack of green tissue above the veins. It can be seen in some aroids. The blessed milk thistle, Silybum marianum, is a plant in which another type of venal variegation occurs, but in this case it is due to a blister variegation occurring along the veins.
Iron and magnesium deficiencies are common causes of this. Transposable elements can cause colour variegation.
Citrus variegation virus (CVV) is a plant pathogenic virus, a member of subgroup 2 of ilarviruses in the family Bromoviridae, is the causal agent of infectious variegation, a disease occurring all over the world, causing problems for production especially in some susceptible varieties of lemon and mandarin.
It has been suggested that some patterns of leaf variegation may be part of a defensive "masquerade strategy." In this, leaf variegation may appear to an insect leaf miner that the leaf is already infested, and this may reduce parasitization of the leaf by leaf miners.
His responsiveness to never-identical site conditions quite obviously allowed for the variegation that permeates his work.
By convention, the italicised term variegata as the second part of the Latin binomial name, indicates a species found in the wild with variegation (Aloe variegata). The much more common, non-italicised, inclusion of 'Variegata' as the third element of a name indicates a variegated cultivar of an unvariegated parent (Aucuba japonica 'Variegata'). However, not all variegated plants have this Latin tag, for instance many cultivars of Pelargonium have some zonal variegation in their leaves. Other types of variegation may be indicated, e.g.
Pilea (aluminum plant) is an example of a house plant that shows this effect. Leaves of most Cyclamen species show such patterned variegation, varying between plants, but consistent within each plant. Another type of reflective variegation is caused by hairs on parts of the leaf, which may be coloured differently from the leaf. This is found in various Begonia species and garden hybrids.
An ornamental plant, F. japonica 'Spider's Web' (or 'Spider White') is a rare cultivar with variegated leaves. Slower growing than the original species, it reaches a lower maximum height of at maturity. The dark-green leaves are strongly white-flecked, particularly at the edges, though the white variegation may occasionally disperse across the whole leaf. The variegation may change with the seasons and as the plant ages.
Variegation of fruits and wood in Ficus carica 'Panascè', a bicolor (yellow- green) common fig cultivar. This Italian cultivar is a chimera Variegation is the appearance of differently coloured zones in the leaves and sometimes the stems, of plants. Variegated leaves occur rarely in nature. Species with variegated individuals are sometimes found in the understory of tropical rainforests, and this habitat is the source of a number of variegated house plants.
Leaves of 300x300px A common cause of variegation is the masking of green pigment by other pigments, such as anthocyanins. This often extends to the whole leaf, causing it to be reddish or purplish. On some plants however, consistent zonal markings occur; such as on some clovers, bromeliads, certain Pelargonium and Oxalis species. On others, such as the commonly grown forms of Coleus, the variegation can vary widely within a population.
SET (Suppressor of variegation, Enhancer of Zeste, Trithorax) and MYND (myeloid-Nervy-DEAF-1) domain-containing protein 3 is a protein that in humans is encoded by the SMYD3 gene.
The forewings have green and blackish variegation. A diffuse sub-dorsal white patch can be seen from the distal end to the postmedial. Transverse fasciae are pale green and irregular.
'Flight's Fancy' is distinguished by its yellow-variegated foliage, the variegation occurring as small blotches across the entirety of the leaf surface. The cultivar is otherwise identical to Buddleja × weyeriana 'Sungold'.
Dieffenbachia seguine is cultivated as an ornamental plant in temperate shade gardens and as a potted house plant. Cultivars emphasize different patterns of variegation. Dieffenbachia seguine grown as a house plant.
Michaelis worked primarily on the inheritance and segregation of cytoplasmic mutations in Epilobium, a genus related to Oenothera. He calculated in 1955/1956 the statistical probability of the segregation of two different types of plastids under different conditions. With further research Michaelis concluded that segregation of a mutant plastid could be the only cause of the observed variegation in the species. After this Michaelis examined thirty cases of plastid variegation where no mixed cells were found.
Since variegation is produced through rare somatic mutations in the apical meristem, this is the preferred method for generating variegated specimens due to the volume of leaves which can be rooted simultaneously.
Numerous cultivars have been developed for garden use, often with variegated foliage. The cultivar 'Argenteovariegatum', with cream-white variegation on the leaves, has gained the Royal Horticultural Society's Award of Garden Merit.
Begonia brevirimosa is a species of flowering plant in the family Begoniaceae. It is native to New Guinea. The pink variegation occurs naturally. It grows as a plant up to 3 m tall.
Wesmael's herbarium specimen shows a young ‘Lutescens’-type leaf with mixed variegation and green (see ‘External links’). Klehm's Ulmus americana variegata had small leaves sprinkled over with white spots, the variation being described as constant.
The other model is the variegation model. Variegated landscapes retain much of their natural vegetation but are intermixed with gradients of modified habitat This model of habitat fragmentation typically applies to landscapes that are modified by agriculture. In contrast to the fragmentation model that is denoted by isolated patches of habitat surrounded by unsuitable landscape environments, the variegation model applies to landscapes modified by agriculture where small patches of habitat remain near the remnant original habitat. In between these patches are a matrix of grassland that is often modified versions of the original habitat.
Receiving a generally favourable reception, AllMusic praising the record an example of the band's best work, Angry Cyclist included politically-angled songs reflecting Brexit and Donald Trump, and absorbed a variegation of influences including country, jangle pop and soul.
HP1 is also found at euchromatic sites, where its binding correlates with gene repression. HP1 was originally discovered by Tharappel C James and Sarah Elgin in 1986 as a factor in the phenomenon known as position effect variegation in Drosophila melanogaster.
It is a somewhat difficult maple to propagate, and is rarely seen outside of sizable collections. Examples may be seen in Westonbirt Arboretum (Gloucestershire, England) and the Arnold Arboretum (Boston, USA). The rare cultivars 'Me uri no ofu' and 'Veitchii' both have striking white and pink variegation.
Here it is also possible to find elements of gypsy scales which contain augmented intervals unusual for the traditional music of Western Europe.Plocek, p. 40 One of the most important elements of the traditional music of Southeastern Moravia is emotional variegation and greater rhythmic leeway.Plocek, p.
Several interesting cultivars have been developed, many of these bear Japanese names. Notable cultivars include 'Goshiki Kaede' (striking pink and green variegation), 'Kifu Nishiki' (roundish, almost un-lobed leaves), 'Mino Yatsubusa' (dwarf with long, narrow leaves) 'Mitsubato Kaede' (distinctive cork-like trunk) and 'Naruto' (strongly incurved leaf surface).
These traces occur in three distinct alluvial depositional settings: flood basin/alluvial plain, crevasse splay, and fluvial channel. Flood basin deposits (dominated by alluvial paleosols with pronounced color variegation) are characterized by common Planolites, rare Skolithos and small, meniscate plug-shaped burrows, possibly Celliforma.Zonneveld et al., 2006, p.
It proceeds to invade the cortical tissue in the stem. In leaves, the rust causes chlorosis and variegation, which might be surrounded by anthocyanescence. One sign of an infection is red-orange filamentous growth emerging on wounds in humid conditions. The most extreme symptoms produce necrotic patches on the stem.
It is likely, however, that 'Pulverulenta' was the U. 'Viminalis Variegata', Variegated Twiggy-branched elm, that was listed and described by John Frederick Wood, F.H.S., in The Midland Florist and Suburban Horticulturist 1847 and 1851, pre-dating both Kirchner and Dieck. Wood did not specify the nature of the variegation.
The tree has been described as a form of 'Pendula' (: 'Horizontalis') with beautiful white-variegated leaves. Pontey (1850) described 'Pendula Variegata' as "distinctly striped and margined with silver" and "remarkable for its constancy in variegation", Wood (1851) as "a first rate ornamental tree" with "beautifully striped foliage" and pendulous branches.
The 'Centennial Variegated' kumquat cultivar arose spontaneously from the oval kumquat. It produces a greater proportion of fruit to peel than the oval kumquat, and the fruit are rounder and sometimes necked. Fruit are distinguishable by their variegation in color, exhibiting bright green and yellow stripes, and by its lack of thorns.
Some mining insects feed in other parts of a plant, such as the surface of a fruit. It has been suggested that some patterns of leaf variegation may be part of a defensive strategy employed by plants to deceive adult leaf miners into thinking that a leaf has already been preyed upon.
There are many proposed models for how the genes in these regions are expressed, including the insulation, denial, integration, exploitation, and TE restraining models. When genes are placed near a region of constitutive heterochromatin, their transcription is usually silenced. This is known as position-effect variegation and can lead to a mosaic phenotype.
Depending on conditions, a plantlet can take anywhere from a few weeks to almost a year before producing new roots, and may take even longer to grow fine roots and become established. Division is the preferred method for propagating most specimens, and in particular, variegated specimens, since leaf cuttings typically do not preserve the variegation.
In wild, the leaves are dark green and without variegation. Cultivated varieties have leaves in various shades of green, often light green and usually with different types of lighter tannins. There are several variegated cultivars, the main differences being in the position and extent of the cream or white markings. Some leaves are almost entirely white, pink or yellow.
Virus infections may cause patterning to appear on the leaf surface. The patterning is often characteristic of the infection. Examples are the mosaic viruses, which produce a mosaic-type effect on the leaf surface or the citrus variegation virus (CVV). Recently a virus disease, Hosta Virus X (HVX) has been identified that causes mottled leaf coloring in hostas.
It has dark green, variegated leaves in length, and in width. The variegation of the leaves arises from the distinct white veins contrasted with the dark green of the leaf. The stems emerge from creeping rhizomes, growing tall. The nearly round flowers, which appear in late July to early August, are found on top of tall stalks.
Numerous cultivars are available (e.g. 'Danielle', 'Naomi', 'Exotica', and 'Golden King'). Some cultivars include different patterns of colouration on the leaves, ranging from light green to dark green, and various forms of white variegation. In cultivation in the UK, this plant and the variegated cultivar 'Starlight' have gained the Royal Horticultural Society's Award of Garden Merit.
This gene is a member of the suppressor of variegation 3-9 homolog family and encodes a protein with a chromodomain and a C-terminal SET domain. This nuclear protein moves to the centromeres during mitosis and functions as a histone methyltransferase, methylating Lys-9 of histone H3. Overall, it plays a vital role in heterochromatin organization, chromosome segregation, and mitotic progression.
Further, he argued, the stereotypical denigration based on race was a matter of early indoctrination, had economic ends, and was supported by the way white clergy condoned slavery.Rita Roberts, Evangelicalism and the Politics of Reform in Northern Black Thought, 1776-1863 (2011), p. 134; Google Books. Easton argued for race as no more intrinsic than any other effect of variegation.
The second section, "Theology", is the most significant. It discusses Iolo's elaborate philosophy and cosmology, which he claimed was the actual belief system of the ancient bards, though it is really a variegation of Iolo's own unconventional Christian beliefs and interpretation of Welsh tradition.Matthews, p. xvi. This section includes a "catechism" in the form of a question and answer session regarding the cycle of re-incarnation.
Aucuba japonica are dioecious. The flowers are small, diameter, each with four purplish-brown petals; they are produced in clusters of 10-30 in a loose cyme. The fruit is a red drupe approximately in diameter, which is avoided by birds. The variegation, considered by some to be an attractive property, is caused by 'Aucuba bacilliform', a putative species of virus in the genus Badnavirus.
There is a spine on each side of the 6th and 7th segments at the apex, the 7th having a third thin spine in the middle. The legs exhibit variegation of yellow and are covered with grey hairs. Female A. manicatum are smaller in size than the males, but have a similar color pattern. The abdominal spots are smaller and the apex is rounded.
The diseased plants are ornamentally desired. The mosaic symptom is considered more interesting than the healthy plant, and it is often marketed as a form of variegation. This disease was one of the first viruses described in scientific literature due to its ornamental properties. Horticulturalists were very interested in the diseases ability to create a variegated look without having to wait for a genetic mutation to pop up.
Leaves become mottled with white or yellow mosaic that resembles variegation. The mottled patches are angular in appearance, and they are limited by veins. Some plants experience stunting, a decrease in photosynthetic potential caused by the reduction of chlorophyll, and they can dehydrate more readily. The symptoms are cosmetic in nature, so infection does not kill the plant, which will still be able to flower and reproduce even when infected.
The first method has the disadvantage that the variegation will be lost. D. trifasciata is considered by some authorities as a potential weed in Australia, although widely used as an ornamental, in both the tropics outdoors in both pots and garden beds and as an indoor plant in temperate areas. The plant contains saponins which are mildly toxic to dogs and cats and can lead to gastrointestinal upset if consumed.
Variations cause heterochromatin to encroach on adjacent genes or recede from genes at the extremes of domains. Transcribable material may be repressed by being positioned (in cis) at these boundary domains. This gives rise to expression levels that vary from cell to cell, which may be demonstrated by position-effect variegation. Insulator sequences may act as a barrier in rare cases where constitutive heterochromatin and highly active genes are juxtaposed (e.g.
Mandiner identifies itself as national liberal and national conservative. It also uses the word "szabadelvű" to describe its line, which is a special Hungarian version of liberalism stemming from the 19th century. "We are not independent, but we also look at ourselves and the political camp close to us with irony. We believe in the variegation of opinions, freedom, tradition and Hungarian history..." - Mandiner wrote on its Facebook page.
This enzyme utilizes a catalytically active site called the SET domain (Suppressor of variegation, Enhancer of zeste, Trithorax). The SET domain is a 130-amino acid sequence involved in modulating gene activities. This domain has been demonstrated to bind to the histone tail and causes the methylation of the histone. Differing histone modifications are likely to function in differing ways; acetylation at one position is likely to function differently from acetylation at another position.
Plants selected by Glenn are grown throughout the world. The patented Euphorbia x martini 'Ascot Rainbow' is to be found in gardens throughout Europe and the United States where it is grown for its frost tolerance and intense variegation which is said to be quite unmatched in the Genus.'Qantas: The Australian Way', #236, p. 68 The plant featured on the cover of the Royal Horticultural Society journal 'The Plantsman' in March 2013.
Position effect is the effect on the expression of a gene when its location in a chromosome is changed, often by translocation. This has been well described in Drosophila with respect to eye color and is known as position effect variegation (PEV). The phenotype is well characterised by unstable expression of a gene that results in the red eye coloration. In the mutant flies the eyes typically have a mottled appearance of white and red sectors.
It is a common houseplant in temperate climates for its striking variegated leaves and purple underside of its leaves. It can grow outside in a humid tropical climate, but needs light shade in the afternoon and must be protected from high winds. The soil should be kept moist at all times, but never waterlogged as they are susceptible to root rot. The distinctive variegation is lost when the plant is under full shade as the leaves become solid green.
Myeloid/lymphoid or mixed-lineage leukemia 4, also known as MLL4, is a human gene. This gene encodes a protein which contains multiple domains including a CXXC zinc finger, three PHD zinc fingers, two FY-rich domains, and a SET (suppressor of variegation, enhancer of zeste, and trithorax) domain. The SET domain is a conserved C-terminal domain that characterizes proteins of the MLL (mixed-lineage leukemia) family. This gene is ubiquitously expressed in adult tissues.
A variant of apple mint, pineapple mint, displays white variegation on its leaves, Pierce County, Washington Mentha suaveolens, the apple mint, pineapple mint, woolly mint or round-leafed mint (synonyms M. rotundifolia, Mentha macrostachya, Mentha insularis), is a member of the mint family Lamiaceae. It is native to southern and western Europe including the Mediterranean region. It is a herbaceous, upright perennial plant that is most commonly grown as a culinary herb or for ground cover.
The band performed in the UK in April 1982. NME considered that the show "was all very skillfully deployed: a bruisingly clear sound of scathing force, a clockwork, Pavlovian lightshow, a variegation of light and shade in the song order that builds to the unmitigating force of 'Pornography' itself as the climax". However, the mood on stage was not good: The journalist noted that Smith looked "dejected and tired" for his birthday. Behind the scene, Smith's relationship with Gallup was deteriorating.
At the Dominion Arboretum, Ottawa, the tree was listed as U. procera 'Marginata', as the variegation is sometimes most obvious on leaf- margins.Photograph of 'Argenteo-Variegata' leaves in Gerald Wilkinson, Epitaph for the Elm, Hutchinson, London 1978, p.67 ( / 0-09-921280-3) Variegated English elm is not to be confused with the more common Field Elm cultivar U. minor 'Argenteo-Variegata', also known as U. minor 'Variegata', the Silver Elm or Tartan Elm, which has similar markings but narrower leaves.
There are a number of closely related taxa which hybridize with L. galeobdolon and in some cases are not unequivocally accepted as distinct species but considered subspecies or varieties by many authors. Most well-known among these is variegated yellow archangel (subsp. argentatum), whose leaves often have variegation, showing as silver patches arranged as a wide semicircle. This, and in particular its large-flowered and even stronger-marked cultivar 'Variegatum', is the taxon most often met with as a garden escapee.
It has variegated leaves of green with pale yellow variegation parallel to the veins. Nurserymen folk legend has it that in 1923, or thereabouts, a consignment of Canna 'Trinacria' rhizomes was despatched to Siam, now Thailand, from a nursery in California. When it arrived at its destination and was grown out it was found to have variegated foliage. A sample was returned to the nursery with a demand for a refund or replacement as it was not what had been ordered.
This is a very variable species with the wing colour ranging from whitish through buff and brown to black, sometimes with variegation, but it is always easily identifiable by the white discal spot on each wing and the characteristic "ragged" margin of the wings. The adults fly at night in May and June and are attracted to light. The caterpillar is green or brown and twig like, and feeds on a variety of plants (see list below). The species overwinters as a pupa.
From this evidence, McClintock hypothesized that there must be a structure on the chromosome tip that would normally ensure stability. She showed that the loss of ring-chromosomes at meiosis caused variegation in maize foliage in generations subsequent to irradiation resulting from chromosomal deletion. During this period, she demonstrated the presence of the nucleolus organizer region on a region on maize chromosome 6, which is required for the assembly of the nucleolus. In 1933, she established that cells can be damaged when nonhomologous recombination occurs.
Goodyera pubescens, the downy rattlesnake plantain, (also known as Peramium pubescens ) is one of the most common orchids native to eastern North America. It is found from Florida to Nova Scotia, west to eastern Oklahoma, Minnesota and Ontario.Kew World Checklist of Selected Plant FamiliesBiota of North America Program county distribution map Goodyera pubescens is an evergreen terrestrial herb with variegated leaves. The variegation is in the form of a densely reticulated network of veins that are a much lighter green than the rest of the leaf tissue.
Neural mapThe shell has a classic form, with smooth shell and edges, unlike Pecten maximus (common name the "great scallop" or "king scallop") which has flutes and scalloped edges; size is around 80 mm, with individuals up to 170 mm in diameter. The shell is generally pinkish-red in color, with striped variegation to darker shades appearing in many individuals. The adductor muscle itself is large, often 30–40 mm in diameter. Like all scallops, P. magellanicus has photoreceptive eyes along the edge of the pinkish mantle.
Many understory plants such as the saw greenbriar, Smilax bona-nox are variegated with pale markings which may serve as camouflage. The possibility of protective coloration in plants has been little studied. T. J. Givnish and Simcha Lev-Yadun have proposed that leaf variegation with white spots may serve as camouflage in forest understory plants, where there is a dappled background. Lev-Yadun has also suggested, however, that similar markings serve as conspicuous warning coloration in well-defended thorny plants of open habitats, where the background is uniformly bright.
316x316px In temperate regions it is a popular houseplant with numerous cultivars selected for leaves with white, yellow, or light green variegation. It is often used in decorative displays in shopping centers, offices, and other public locations largely because it requires little care and is also attractively leafy. In tropical countries, it is found in many parks and gardens and tends to grow naturally. As an indoor plant it can reach more than 2 m in height if it is given the adequate support (a tutor to climb), but hardly develops adult-sized leaves.
With a minimum temperature of , it must be grown indoors in most temperate regions. Although it needs high humidity, it does not require much care, it is enough with good lighting (without becoming excessive) to maintain the bright and intense colour of the leaves, these together with the stem are fleshy, so they do not need a very abundant or frequent watering. Direct light causes discoloration of its leaves, its average lighting is enough. A popular houseplant, numerous cultivars have been developed, some of which show leaf variegation.
In the anaphase of mitosis, the broken chromosomes formed a chromatid bridge, which was broken when the chromatids moved towards the cell poles. The broken ends were rejoined in the interphase of the next mitosis, and the cycle was repeated, causing massive mutation, which she could detect as variegation in the endosperm. This breakage–rejoining–bridge cycle was a key cytogenetic discovery for several reasons. First, it showed that the rejoining of chromosomes was not a random event, and second, it demonstrated a source of large-scale mutation.
Maize streak virus (MSV) causes a plant disease, known as maize streak disease (MSD) in its major host. It is an insect-transmitted maize pathogen in the genus Mastrevirus of the family Geminiviridae that is endemic in sub-Saharan Africa and neighbouring Indian Ocean island territories such as Madagascar, Mauritius and La Reunion. The A-strain of MSV (MSV-A) causes sporadic maize streak disease epidemics throughout the maize growing regions of Africa. MSV was first described by the South African entomologist Claude Fuller who referred to it in a 1901 report as "mealie variegation".
The majority of variegated-leaf chimeras are of this kind. All plastid gene- and some nuclear gene-differential chimeras affect the color of the plasmids within the leaves, and these are grouped together as chlorophyll chimeras, or preferably as variegated leaf chimeras. For most variegation, the mutation involved is the loss of the chloroplasts in the mutated tissue, so that part of the plant tissue has no green pigment and no photosynthetic ability. This mutated tissue is unable to survive on its own, but it is kept alive by its partnership with normal photosynthetic tissue.
Plocek, p. 40 Distinct styles of folk music began to emerge during the Wallachian colonization of the 16th and 17th centuries, separating Bohemian and Moravian traditional music. The "new Hungarian" style has influenced the music of the area in the past three centuries, especially in Southern Moravia.Plocek, p. 40 This influence has left a deep imprint on the unusual melodic variegation of Moravian traditional music providing an inspirational source for subsequent Classical, Jazz and Pop music composers.Plocek, p. 40 The Czech National Revival in the 19th century represented an important turning point for traditional music.
Actinidia kolomikta is an ornamental plant for gardens and a houseplant. The plant was collected by Charles Maries in Sapporo, on the northern Japanese island of Hokkaido, in 1878, and sent to his patrons, Veitch Nurseries, who introduced it into Western horticulture.Alice M. Coats, Garden Shrubs and Their History (1964) 1992, s.v. "Actinidia". Actinidia kolomikta is cultivated in cold temperate regions as an ornamental plant, largely for the striking random variegation in pink and white of some its leaves but also because of the relatively small (2-5 g or 0.07- 0.18 ounces) kiwifruit-like berries it produces.
His main focuses were phytopathology, cecidogenesis (the forming of galls), non-pathogenic cancer growth, non-pathogenic nodule growth, facultative and obligatory nodosities (root deformations), mycotic causes of wilting of some fruit tree species, and the problem of early death of apricot trees. He was a knowledgeable expert in the research field of flaws of timber, especially the red heartwood core of beeches and the variegation of beech wood. Throughout his life, Paclt worked on various groups of hexapods. His main groups of interest were springtails, proturans, diplurans, jumping bristletails, silverfish, butterflies and moths, beetles and mayflies, of which he studied the morphology, taxonomy, systematics, ecology and zoogeography.
The virus infects the bulb and causes the cultivar to "break" its lock on a single color, resulting in intricate bars, stripes, streaks, featherings or flame-like effects of different colors on the petals. These symptoms vary depending on the plant variety and age at the time of infection. Different types of colour-breaks depend on the variety of tulip and the strain of the virus. The color variegation is caused either by local fading, or intensification and overaccumulation of pigments in the vacuoles of the upper epidermal layer due to the irregular distribution of anthocyanin; this fluctuation in pigmentation occurs after the normal flower color has developed.
France sits at the edge of the European peninsula and has seen waves of migration of groups that often settled owing to the presence of physical barriers preventing onward migration. This has led to language and regional cultural variegation, but the extent to which this pattern of migrations showed up in population genetics studies was unclear until the publication of a study in 2019 that used genome wide data. The study identified six different genetic clusters that could be distinguished across populations. The study concluded that the population genetic clusters correlate with linguistic and historical divisions in France and with the presence of geographic barriers such as mountains and major rivers.
Laura Riding, in her poem The Map of Places (1927), deals with this relation: "The map of places passes. The reality of paper tears." The economist Joan Robinson (1962): "A model which took account of all the variegation of reality would be of no more use than a map at the scale of one to one." Korzybski's argument about the map and the territory also influenced the Belgian surrealist writer of comics Jan Bucquoy for a storyline in his comic Labyrinthe: a map can never guarantee that one will find the way out, because the accumulation of events can change the way one looks at reality.
Variegation produced by the tulip breaking virus Botrytis tulipae is a major fungal disease affecting tulips, causing cell death and eventually the rotting of the plant. Other pathogens include anthracnose, bacterial soft rot, blight caused by Sclerotium rolfsii, bulb nematodes, other rots including blue molds, black molds and mushy rot. The fungus Trichoderma viride can infect tulips, producing dried leaf tips and reduced growth, although symptoms are usually mild and only present on bulbs growing in glasshouses. Variegated tulips admired during the Dutch tulipomania gained their delicately feathered patterns from an infection with the tulip breaking virus, a mosaic virus that was carried by the green peach aphid, Myzus persicae.
Its distinctive features are the pronounced constriction in the back of the head (as if the waist were in the back of the head), a shallow furrow along the middle of the back from between the eyes to near the sacral hump, and the short, rounded snout. Other distinguishing but not necessarily exclusive characteristics are: the small eyes, with narrow upper eyelids, the uniformerly granular dorsum, the pair of light, externally concave lines on the back (not too distinct in very dark animals), the blackish throat of males, and the absence of dark streaks along the sides of the snout. The basic dorsal color may be brown or dark gray, almost uniform or with obscure variegation or vermiculations of lighter gray. The males average 32.8 mm in length, while females average 34.6 mm.
Gouache on paper drawing, before 1640, of the Semper Augustus, famous for being the most expensive tulip sold during tulip mania. The effects of Tulip breaking virus are seen in the striking streaks of white in its red petals. Long thought to be the earliest recorded plant virus, it is now thought that TBV comes second; the earliest reference to a virus-induced leaf chlorosis (possibly tobacco leaf curl virus) was recorded in Japan in 752 AD.Brunt, Alan; Walsh, John, "'Broken' tulips and Tulip breaking virus", Microbiology Today, May, 2005, p. 71. "Breaking" symptomology was first described in 1576 by Carolus Clusius, a Flemish professor of Botany at Leiden, who noted the variegation, or "rectification", so termed because it was believed that with the offset production of an entirely new "broken" bloom the plant was distilling, or rectifying, itself into a pure life form.

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